Topography as a determinant of range extent and overlap : a species level phylogenetic reconstruction and geographical range analysis of Syncarpha (Asteraceae)

dc.contributor.advisorVerboom, George Anthonyen_ZA
dc.contributor.advisorBergh, Nicola Gen_ZA
dc.contributor.authorHaiden, Sarahen_ZA
dc.date.accessioned2017-11-10T09:29:18Z
dc.date.available2017-11-10T09:29:18Z
dc.date.issued2011en_ZA
dc.date.updated2017-03-13T10:34:32Z
dc.description.abstractUnderstanding what determines species' geographic range extents has several implications for questions in ecology, evolution and conservation biology. The Cape Floristic Region of South Africa is noted for its remarkably high geographic species turnover, often attributed to the exceptional environmental heterogeneity of the region. The complex and highly dissected topography of the CFR provides a model environment in which to investigate the relationship between altitude and species range extent, as well as explore the role of topography in speciation and current range overlap. I examined these questions in the context of Syncarpha, a genus within the Asteraceae tribe Gnaphalieae (paper daisies). A Bayesian analysis of combined plastid and nuclear genes provided the robust, dated phylogenetic hypothesis required to assess the monophyly of the genus, as well as reconstruct the signal of geographic speciation within the lineage. The phylogeny recovered Syncarpha as polyphyletic, comprising two clades with good support, placing the small CFR-endemic genus Edmondia as sister to the larger Syncarpha clade. Using realised range extent estimates and modelled potential distributions of Syncarpha and Edmondia species, this study confirms the importance of topography as a factor constraining species' distributions, and thereby enhancing the scope for their allopatric isolation. The relationship between altitude and realised range extent was found to be unimodal, with ranges being restricted at both high- and low-altitudes, and more extensive at intermediate altitudes. Range filling (the ratio between realised and potential range extent) was also lower in high- and low-altitude taxa compared to mid-altitude taxa. Dispersal limitation, owing to the insular nature of montane habitats, seems the most likely mechanism to restrict the ranges of high-altitude taxa, whereas edaphic factors are more likely responsible for the restricted ranges of low-altitude taxa. Furthermore, age-range correlations confirm the role of altitude in maintaining a stronger signal of allopatry among recently diverged clades, where montane clades present lower levels of range overlap than those at intermediate altitudes. Thus, the role of topography in limiting dispersal, and hence constraining species distributions, has consequences for understanding the historical diversification of a lineage, as well as implications for management practices in light of climate change-induced range shifts.en_ZA
dc.identifier.apacitationHaiden, S. (2011). <i>Topography as a determinant of range extent and overlap : a species level phylogenetic reconstruction and geographical range analysis of Syncarpha (Asteraceae)</i>. (Thesis). University of Cape Town ,Faculty of Science ,Department of Biological Sciences. Retrieved from http://hdl.handle.net/11427/26142en_ZA
dc.identifier.chicagocitationHaiden, Sarah. <i>"Topography as a determinant of range extent and overlap : a species level phylogenetic reconstruction and geographical range analysis of Syncarpha (Asteraceae)."</i> Thesis., University of Cape Town ,Faculty of Science ,Department of Biological Sciences, 2011. http://hdl.handle.net/11427/26142en_ZA
dc.identifier.citationHaiden, S. 2011. Topography as a determinant of range extent and overlap : a species level phylogenetic reconstruction and geographical range analysis of Syncarpha (Asteraceae). University of Cape Town.en_ZA
dc.identifier.ris TY - Thesis / Dissertation AU - Haiden, Sarah AB - Understanding what determines species' geographic range extents has several implications for questions in ecology, evolution and conservation biology. The Cape Floristic Region of South Africa is noted for its remarkably high geographic species turnover, often attributed to the exceptional environmental heterogeneity of the region. The complex and highly dissected topography of the CFR provides a model environment in which to investigate the relationship between altitude and species range extent, as well as explore the role of topography in speciation and current range overlap. I examined these questions in the context of Syncarpha, a genus within the Asteraceae tribe Gnaphalieae (paper daisies). A Bayesian analysis of combined plastid and nuclear genes provided the robust, dated phylogenetic hypothesis required to assess the monophyly of the genus, as well as reconstruct the signal of geographic speciation within the lineage. The phylogeny recovered Syncarpha as polyphyletic, comprising two clades with good support, placing the small CFR-endemic genus Edmondia as sister to the larger Syncarpha clade. Using realised range extent estimates and modelled potential distributions of Syncarpha and Edmondia species, this study confirms the importance of topography as a factor constraining species' distributions, and thereby enhancing the scope for their allopatric isolation. The relationship between altitude and realised range extent was found to be unimodal, with ranges being restricted at both high- and low-altitudes, and more extensive at intermediate altitudes. Range filling (the ratio between realised and potential range extent) was also lower in high- and low-altitude taxa compared to mid-altitude taxa. Dispersal limitation, owing to the insular nature of montane habitats, seems the most likely mechanism to restrict the ranges of high-altitude taxa, whereas edaphic factors are more likely responsible for the restricted ranges of low-altitude taxa. Furthermore, age-range correlations confirm the role of altitude in maintaining a stronger signal of allopatry among recently diverged clades, where montane clades present lower levels of range overlap than those at intermediate altitudes. Thus, the role of topography in limiting dispersal, and hence constraining species distributions, has consequences for understanding the historical diversification of a lineage, as well as implications for management practices in light of climate change-induced range shifts. DA - 2011 DB - OpenUCT DP - University of Cape Town LK - https://open.uct.ac.za PB - University of Cape Town PY - 2011 T1 - Topography as a determinant of range extent and overlap : a species level phylogenetic reconstruction and geographical range analysis of Syncarpha (Asteraceae) TI - Topography as a determinant of range extent and overlap : a species level phylogenetic reconstruction and geographical range analysis of Syncarpha (Asteraceae) UR - http://hdl.handle.net/11427/26142 ER - en_ZA
dc.identifier.urihttp://hdl.handle.net/11427/26142
dc.identifier.vancouvercitationHaiden S. Topography as a determinant of range extent and overlap : a species level phylogenetic reconstruction and geographical range analysis of Syncarpha (Asteraceae). [Thesis]. University of Cape Town ,Faculty of Science ,Department of Biological Sciences, 2011 [cited yyyy month dd]. Available from: http://hdl.handle.net/11427/26142en_ZA
dc.language.isoengen_ZA
dc.publisher.departmentDepartment of Biological Sciencesen_ZA
dc.publisher.facultyFaculty of Scienceen_ZA
dc.publisher.institutionUniversity of Cape Town
dc.subject.otherBotanyen_ZA
dc.titleTopography as a determinant of range extent and overlap : a species level phylogenetic reconstruction and geographical range analysis of Syncarpha (Asteraceae)en_ZA
dc.typeBachelor Thesis
dc.type.qualificationlevelHonours
dc.type.qualificationnameBSc (Hons)en_ZA
uct.type.filetype
uct.type.filetypeText
uct.type.filetypeImage
uct.type.publicationResearchen_ZA
uct.type.resourceThesisen_ZA
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