Coevolutionary causes and consequences of high-fidelity mimicry by a specialist brood parasite

dc.contributor.advisorSpottiswoode, Claire N
dc.contributor.advisorJamie, Gabriel A
dc.contributor.authorLund, Jess
dc.date.accessioned2022-03-04T08:05:17Z
dc.date.available2022-03-04T08:05:17Z
dc.date.issued2021
dc.date.updated2022-03-03T13:11:45Z
dc.description.abstractMimicry is often invoked as a classic demonstration of the power of natural selection. However, mimicry systems are diverse and the accuracy of mimicry, or mimetic fidelity, varies from crude to near-perfect. There is growing evidence that low-fidelity mimicry is the norm, with cases of high-fidelity mimicry being rarer. This is particularly evident in avian interspecific brood parasites, which are birds that lay their eggs in the nests of other species, or ‘hosts'. A possible rare example of a “perfect mimic” is the African cuckoo Cuculus gularis (a specialist parasite of fork-tailed drongos Dicrurus adsimilis) which, to the human eye, exhibits some of the most sophisticated mimicry seen in a brood parasite-host system. In this work, I quantify the degree of mimetic fidelity in this system and investigate its consequences on the two antagonists, using experimental, observational and genetic data I collected in the field in Zambia. In Chapter 1 I first define perfect mimicry, and explore how mimetic fidelity can be quantified. I then review the factors responsible for the variability in mimetic fidelity among different mimicry systems. In Chapter 2 I use quantitative measures of egg colour and pattern, from models that approximate avian vision, to show that mimicry of drongo eggs by African cuckoos is near-perfect. I subsequently use field experiments and simulations to show that a high degree of interclutch variation (egg ‘signatures') means that drongos still have the upper hand in the arms race against this near-perfect mimic. In Chapter 3 I investigate the genetic architecture of perfect mimicry by comparing cuckoo egg phenotypes to their mitochondrial DNA haplotypes, providing evidence that the African cuckoo likely shows bi-parental inheritance of egg phenotype. Together these results highlight the effectiveness of egg signatures as a defence against brood parasites, even when mimicry shows high accuracy and precision, and underline the role of mechanistic factors in enabling high-fidelity mimicry to evolve.
dc.identifier.apacitationLund, J. (2021). <i>Coevolutionary causes and consequences of high-fidelity mimicry by a specialist brood parasite</i>. (). ,Faculty of Science ,Department of Biological Sciences. Retrieved from http://hdl.handle.net/11427/35903en_ZA
dc.identifier.chicagocitationLund, Jess. <i>"Coevolutionary causes and consequences of high-fidelity mimicry by a specialist brood parasite."</i> ., ,Faculty of Science ,Department of Biological Sciences, 2021. http://hdl.handle.net/11427/35903en_ZA
dc.identifier.citationLund, J. 2021. Coevolutionary causes and consequences of high-fidelity mimicry by a specialist brood parasite. . ,Faculty of Science ,Department of Biological Sciences. http://hdl.handle.net/11427/35903en_ZA
dc.identifier.ris TY - Master Thesis AU - Lund, Jess AB - Mimicry is often invoked as a classic demonstration of the power of natural selection. However, mimicry systems are diverse and the accuracy of mimicry, or mimetic fidelity, varies from crude to near-perfect. There is growing evidence that low-fidelity mimicry is the norm, with cases of high-fidelity mimicry being rarer. This is particularly evident in avian interspecific brood parasites, which are birds that lay their eggs in the nests of other species, or ‘hosts'. A possible rare example of a “perfect mimic” is the African cuckoo Cuculus gularis (a specialist parasite of fork-tailed drongos Dicrurus adsimilis) which, to the human eye, exhibits some of the most sophisticated mimicry seen in a brood parasite-host system. In this work, I quantify the degree of mimetic fidelity in this system and investigate its consequences on the two antagonists, using experimental, observational and genetic data I collected in the field in Zambia. In Chapter 1 I first define perfect mimicry, and explore how mimetic fidelity can be quantified. I then review the factors responsible for the variability in mimetic fidelity among different mimicry systems. In Chapter 2 I use quantitative measures of egg colour and pattern, from models that approximate avian vision, to show that mimicry of drongo eggs by African cuckoos is near-perfect. I subsequently use field experiments and simulations to show that a high degree of interclutch variation (egg ‘signatures') means that drongos still have the upper hand in the arms race against this near-perfect mimic. In Chapter 3 I investigate the genetic architecture of perfect mimicry by comparing cuckoo egg phenotypes to their mitochondrial DNA haplotypes, providing evidence that the African cuckoo likely shows bi-parental inheritance of egg phenotype. Together these results highlight the effectiveness of egg signatures as a defence against brood parasites, even when mimicry shows high accuracy and precision, and underline the role of mechanistic factors in enabling high-fidelity mimicry to evolve. DA - 2021_ DB - OpenUCT DP - University of Cape Town KW - Biological Sciences LK - https://open.uct.ac.za PY - 2021 T1 - Coevolutionary causes and consequences of high-fidelity mimicry by a specialist brood parasite TI - Coevolutionary causes and consequences of high-fidelity mimicry by a specialist brood parasite UR - http://hdl.handle.net/11427/35903 ER - en_ZA
dc.identifier.urihttp://hdl.handle.net/11427/35903
dc.identifier.vancouvercitationLund J. Coevolutionary causes and consequences of high-fidelity mimicry by a specialist brood parasite. []. ,Faculty of Science ,Department of Biological Sciences, 2021 [cited yyyy month dd]. Available from: http://hdl.handle.net/11427/35903en_ZA
dc.language.rfc3066eng
dc.publisher.departmentDepartment of Biological Sciences
dc.publisher.facultyFaculty of Science
dc.subjectBiological Sciences
dc.titleCoevolutionary causes and consequences of high-fidelity mimicry by a specialist brood parasite
dc.typeMaster Thesis
dc.type.qualificationlevelMasters
dc.type.qualificationlevelMSc
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